Kelly A. Brayton

Conservation in the face of diversity: multistrain analysis of an intracellular bacterium (2009)

Dark, Michael J, Herndon, David R, Kappmeyer, Lowell S, Gonzales, Mikel P, Nordeen, Elizabeth, Palmer, Guy H, ...

Abstract Background With the recent completion of numerous sequenced bacterial genomes, notable advances have been made in understanding the level of conservation between various species. However,...

BMC Genomics BioMed Central (2008)

Michael J Dark, David R Herndon, Mikel P Gonzales, Elizabeth Nordeen, Guy H Palmer, Donald P Knowles, ...

Research article Conservation in the face of diversity: multistrain analysis of an intracellular bacterium

Genome Sequence of Babesia bovis and Comparative Analysis of Apicomplexan Hemoprotozoa (2007)

Kelly A. Brayton, David R. Herndon, Linda Hannick, Lowell S. Kappmeyer, Shawn J. Berens, ...

Babesia bovis is an apicomplexan tick-transmitted pathogen of cattle imposing a global risk and severe constraints to livestock health and economic development. The complete genome sequence was...

Stochastic Transmission of Multiple Genotypically Distinct Anaplasma marginale Strains in a Herd with High Prevalence of Anaplasma Infection (2004)

Guy H. Palmer, Donald P. Knowles, Jose-luis Rodriguez, David P. Gnad, Larry C. Hollis, Twig Marston, ...

Multiple genotypically unique strains of the tick-borne pathogen Anaplasma marginale occur and are transmitted within regions where the organism is endemic. In this study, we tested the hypothesis...

Efficient use of a small genome to generate antigenic diversity in tick-borne ehrlichial pathogens

Brayton, Kelly A., Knowles, Donald P., McGuire, Travis C., Palmer, Guy H.

Ehrlichiae are responsible for important tick-transmitted diseases, including anaplasmosis, the most prevalent tick-borne infection of livestock worldwide, and the emerging human diseases monocytic...

Expression of Major Surface Protein 2 Variants with Conserved T-Cell Epitopes in Anaplasma centrale Vaccinates

Shkap, Varda, Molad, Thea, Brayton, Kelly A., Brown, Wendy C., Palmer, Guy H.

Major surface protein 2 (MSP-2), identified as a protection-inducing immunogen against Anaplasma marginale challenge, is an immunodominant outer membrane protein with orthologues in all examined...

Expression of Anaplasma marginale Major Surface Protein 2 Operon-Associated Proteins during Mammalian and Arthropod Infection

Löhr, Christiane V., Brayton, Kelly A., Shkap, Varda, Molad, Thea, Barbet, Anthony F., Brown, Wendy C., ...

The antigenically variant major surface protein 2 (MSP2) of Anaplasma marginale is expressed from a 3.5-kb operon that contains, in a 5′-to-3′ direction, four open reading frames, opag3, opag2,...

Simultaneous Variation of the Immunodominant Outer Membrane Proteins, MSP2 and MSP3, during Anaplasma marginale Persistence In Vivo

Brayton, Kelly A., Meeus, Patrick F. M., Barbet, Anthony F., Palmer, Guy H.

Vector-borne bacterial pathogens persist in the mammalian host by varying surface antigens to evade the existing immune response. To test whether the model of surface coat switching and immune...

CD4+ T Lymphocytes from Anaplasma marginale Major Surface Protein 2 (MSP2) Vaccinees Recognize Naturally Processed Epitopes Conserved in MSP3

Brown, Wendy C., Palmer, Guy H., Brayton, Kelly A., Meeus, Patrick F. M., Barbet, Anthony F., Kegerreis, Kimberly A., ...

Major surface protein 2 (MSP2) and MSP3 of the persistent bovine ehrlichial pathogen Anaplasma marginale are immunodominant proteins that undergo antigenic variation. The recently completed sequence...

Stochastic Transmission of Multiple Genotypically Distinct Anaplasma marginale Strains in a Herd with High Prevalence of Anaplasma Infection

Palmer, Guy H., Knowles, Donald P., Rodriguez, Jose-Luis, Gnad, David P., Hollis, Larry C., Marston, Twig, ...

Multiple genotypically unique strains of the tick-borne pathogen Anaplasma marginale occur and are transmitted within regions where the organism is endemic. In this study, we tested the hypothesis...

Identification of a Novel Anaplasma marginale Appendage-Associated Protein That Localizes with Actin Filaments during Intraerythrocytic Infection

Stich, Roger W., Olah, Glenn A., Brayton, Kelly A., Brown, Wendy C., Fechheimer, Marcus, Green-Church, Kari, ...

The rickettsial pathogen Anaplasma marginale assembles an actin filament bundle during intracellular infection. Unlike other bacterial pathogens that generate actin filament tails, A. marginale...

The genome of the heartwater agent Ehrlichia ruminantium contains multiple tandem repeats of actively variable copy number

Collins, Nicola E., Liebenberg, Junita, De Villiers, Etienne P., Brayton, Kelly A., Louw, Elmarié, Pretorius, Alri, ...

Heartwater, a tick-borne disease of domestic and wild ruminants, is caused by the intracellular rickettsia Ehrlichia ruminantium (previously known as Cowdria ruminantium). It is a major constraint to...

Complete genome sequencing of Anaplasma marginale reveals that the surface is skewed to two superfamilies of outer membrane proteins

Brayton, Kelly A., Kappmeyer, Lowell S., Herndon, David R., Dark, Michael J., Tibbals, David L., Palmer, Guy H., ...

The rickettsia Anaplasma marginale is the most prevalent tick-borne livestock pathogen worldwide and is a severe constraint to animal health. A. marginale establishes lifelong persistence in infected...

Sequence Variation and Immunologic Cross-Reactivity among Babesia bovis Merozoite Surface Antigen 1 Proteins from Vaccine Strains and Vaccine Breakthrough Isolates

LeRoith, Tanya, Brayton, Kelly A., Molloy, John B., Bock, Russell E., Hines, Stephen A., Lew, Ala E., ...

The Babesia bovis merozoite surface antigen 1 (MSA-1) is an immunodominant membrane glycoprotein that is the target of invasion-blocking antibodies. While antigenic variation has been demonstrated in...

Merozoite Surface Antigen 2 Proteins of Babesia bovis Vaccine Breakthrough Isolates Contain a Unique Hypervariable Region Composed of Degenerate Repeats

Berens, Shawn J., Brayton, Kelly A., Molloy, John B., Bock, Russell E., Lew, Ala E., McElwain, Terry F.

The merozoite surface antigen 2 (MSA-2) proteins of Babesia bovis are members of the variable merozoite surface antigen (VMSA) family that have been implicated in erythrocyte invasion and are...

Identification of Novel Antigenic Proteins in a Complex Anaplasma marginale Outer Membrane Immunogen by Mass Spectrometry and Genomic Mapping

Lopez, Job E., Siems, William F., Palmer, Guy H., Brayton, Kelly A., McGuire, Travis C., Norimine, Junzo, ...

Immunization with purified Anaplasma marginale outer membranes induces complete protection against infection that is associated with CD4+ T-lymphocyte-mediated gamma interferon secretion and...

Differential Expression and Sequence Conservation of the Anaplasma marginale msp2 Gene Superfamily Outer Membrane Proteins

Noh, Susan M., Brayton, Kelly A., Knowles, Donald P., Agnes, Joseph T., Dark, Michael J., Brown, Wendy C., ...

Bacterial pathogens in the genera Anaplasma and Ehrlichia encode a protein superfamily, pfam01617, which includes the predominant outer membrane proteins (OMPs) of each species, major surface protein...

The Babesia bovis Merozoite Surface Antigen 1 Hypervariable Region Induces Surface-Reactive Antibodies That Block Merozoite Invasion

LeRoith, Tanya, Berens, Shawn J., Brayton, Kelly A., Hines, Stephen A., Brown, Wendy C., Norimine, Junzo, ...

A hypervariable region (HVR) previously identified in the carboxy-terminal one-third of the Babesia bovis variable merozoite surface antigen family was more extensively analyzed in merozoite surface...

Analysis of the Anaplasma marginale Major Surface Protein 1 Complex Protein Composition by Tandem Mass Spectrometry

Macmillan, Henriette, Brayton, Kelly A., Palmer, Guy H., McGuire, Travis C., Munske, Gerhard, Siems, William F., ...

The protective major surface protein 1 (MSP1) complex of Anaplasma marginale is a heteromer of MSP1a and MSP1b, encoded by a multigene family. The msp1β sequences were highly conserved throughout...

Efficient use of a small genome to generate antigenic diversity in tick-borne ehrlichial pathogens

Brayton, Kelly A., Knowles, Donald P., McGuire, Travis C., Palmer, Guy H.

Ehrlichiae are responsible for important tick-transmitted diseases, including anaplasmosis, the most prevalent tick-borne infection of livestock worldwide, and the emerging human diseases monocytic...

Expression of Major Surface Protein 2 Variants with Conserved T-Cell Epitopes in Anaplasma centrale Vaccinates

Shkap, Varda, Molad, Thea, Brayton, Kelly A., Brown, Wendy C., Palmer, Guy H.

Major surface protein 2 (MSP-2), identified as a protection-inducing immunogen against Anaplasma marginale challenge, is an immunodominant outer membrane protein with orthologues in all examined...

Expression of Anaplasma marginale Major Surface Protein 2 Operon-Associated Proteins during Mammalian and Arthropod Infection

Löhr, Christiane V., Brayton, Kelly A., Shkap, Varda, Molad, Thea, Barbet, Anthony F., Brown, Wendy C., ...

The antigenically variant major surface protein 2 (MSP2) of Anaplasma marginale is expressed from a 3.5-kb operon that contains, in a 5′-to-3′ direction, four open reading frames, opag3, opag2,...

Simultaneous Variation of the Immunodominant Outer Membrane Proteins, MSP2 and MSP3, during Anaplasma marginale Persistence In Vivo

Brayton, Kelly A., Meeus, Patrick F. M., Barbet, Anthony F., Palmer, Guy H.

Vector-borne bacterial pathogens persist in the mammalian host by varying surface antigens to evade the existing immune response. To test whether the model of surface coat switching and immune...

CD4+ T Lymphocytes from Anaplasma marginale Major Surface Protein 2 (MSP2) Vaccinees Recognize Naturally Processed Epitopes Conserved in MSP3

Brown, Wendy C., Palmer, Guy H., Brayton, Kelly A., Meeus, Patrick F. M., Barbet, Anthony F., Kegerreis, Kimberly A., ...

Major surface protein 2 (MSP2) and MSP3 of the persistent bovine ehrlichial pathogen Anaplasma marginale are immunodominant proteins that undergo antigenic variation. The recently completed sequence...

Stochastic Transmission of Multiple Genotypically Distinct Anaplasma marginale Strains in a Herd with High Prevalence of Anaplasma Infection

Palmer, Guy H., Knowles, Donald P., Rodriguez, Jose-Luis, Gnad, David P., Hollis, Larry C., Marston, Twig, ...

Multiple genotypically unique strains of the tick-borne pathogen Anaplasma marginale occur and are transmitted within regions where the organism is endemic. In this study, we tested the hypothesis...

Identification of a Novel Anaplasma marginale Appendage-Associated Protein That Localizes with Actin Filaments during Intraerythrocytic Infection

Stich, Roger W., Olah, Glenn A., Brayton, Kelly A., Brown, Wendy C., Fechheimer, Marcus, Green-Church, Kari, ...

The rickettsial pathogen Anaplasma marginale assembles an actin filament bundle during intracellular infection. Unlike other bacterial pathogens that generate actin filament tails, A. marginale...

The genome of the heartwater agent Ehrlichia ruminantium contains multiple tandem repeats of actively variable copy number

Collins, Nicola E., Liebenberg, Junita, De Villiers, Etienne P., Brayton, Kelly A., Louw, Elmarié, Pretorius, Alri, ...

Heartwater, a tick-borne disease of domestic and wild ruminants, is caused by the intracellular rickettsia Ehrlichia ruminantium (previously known as Cowdria ruminantium). It is a major constraint to...

Complete genome sequencing of Anaplasma marginale reveals that the surface is skewed to two superfamilies of outer membrane proteins

Brayton, Kelly A., Kappmeyer, Lowell S., Herndon, David R., Dark, Michael J., Tibbals, David L., Palmer, Guy H., ...

The rickettsia Anaplasma marginale is the most prevalent tick-borne livestock pathogen worldwide and is a severe constraint to animal health. A. marginale establishes lifelong persistence in infected...

Sequence Variation and Immunologic Cross-Reactivity among Babesia bovis Merozoite Surface Antigen 1 Proteins from Vaccine Strains and Vaccine Breakthrough Isolates

LeRoith, Tanya, Brayton, Kelly A., Molloy, John B., Bock, Russell E., Hines, Stephen A., Lew, Ala E., ...

The Babesia bovis merozoite surface antigen 1 (MSA-1) is an immunodominant membrane glycoprotein that is the target of invasion-blocking antibodies. While antigenic variation has been demonstrated in...

Merozoite Surface Antigen 2 Proteins of Babesia bovis Vaccine Breakthrough Isolates Contain a Unique Hypervariable Region Composed of Degenerate Repeats

Berens, Shawn J., Brayton, Kelly A., Molloy, John B., Bock, Russell E., Lew, Ala E., McElwain, Terry F.

The merozoite surface antigen 2 (MSA-2) proteins of Babesia bovis are members of the variable merozoite surface antigen (VMSA) family that have been implicated in erythrocyte invasion and are...

Identification of Novel Antigenic Proteins in a Complex Anaplasma marginale Outer Membrane Immunogen by Mass Spectrometry and Genomic Mapping

Lopez, Job E., Siems, William F., Palmer, Guy H., Brayton, Kelly A., McGuire, Travis C., Norimine, Junzo, ...

Immunization with purified Anaplasma marginale outer membranes induces complete protection against infection that is associated with CD4+ T-lymphocyte-mediated gamma interferon secretion and...

Differential Expression and Sequence Conservation of the Anaplasma marginale msp2 Gene Superfamily Outer Membrane Proteins

Noh, Susan M., Brayton, Kelly A., Knowles, Donald P., Agnes, Joseph T., Dark, Michael J., Brown, Wendy C., ...

Bacterial pathogens in the genera Anaplasma and Ehrlichia encode a protein superfamily, pfam01617, which includes the predominant outer membrane proteins (OMPs) of each species, major surface protein...

The Babesia bovis Merozoite Surface Antigen 1 Hypervariable Region Induces Surface-Reactive Antibodies That Block Merozoite Invasion

LeRoith, Tanya, Berens, Shawn J., Brayton, Kelly A., Hines, Stephen A., Brown, Wendy C., Norimine, Junzo, ...

A hypervariable region (HVR) previously identified in the carboxy-terminal one-third of the Babesia bovis variable merozoite surface antigen family was more extensively analyzed in merozoite surface...

Analysis of the Anaplasma marginale Major Surface Protein 1 Complex Protein Composition by Tandem Mass Spectrometry

Macmillan, Henriette, Brayton, Kelly A., Palmer, Guy H., McGuire, Travis C., Munske, Gerhard, Siems, William F., ...

The protective major surface protein 1 (MSP1) complex of Anaplasma marginale is a heteromer of MSP1a and MSP1b, encoded by a multigene family. The msp1β sequences were highly conserved throughout...

Selection for Simple Major Surface Protein 2 Variants during Anaplasma marginale Transmission to Immunologically Naïve Animals▿

Palmer, Guy H., Futse, James E., Leverich, Christina K., Knowles, Donald P., Rurangirwa, Fred R., Brayton, Kelly A.

Anaplasma marginale, a rickettsial pathogen, evades clearance in the animal host by antigenic variation. Under immune selection, A. marginale expresses complex major surface protein 2 mosaics,...

Immunogenicity of Anaplasma marginale Type IV Secretion System Proteins in a Protective Outer Membrane Vaccine▿

Lopez, Job E., Palmer, Guy H., Brayton, Kelly A., Dark, Michael J., Leach, Stephanie E., Brown, Wendy C.

Rickettsial pathogens in the genera Anaplasma and Ehrlichia cause acute infection in immunologically naive hosts and are major causes of tick-borne disease in animals and humans. Immunization with...

Genome Sequence of Babesia bovis and Comparative Analysis of Apicomplexan Hemoprotozoa

Brayton, Kelly A, Lau, Audrey O. T, Herndon, David R, Hannick, Linda, Kappmeyer, Lowell S, Berens, Shawn J, ...

Babesia bovis is an apicomplexan tick-transmitted pathogen of cattle imposing a global risk and severe constraints to livestock health and economic development. The complete genome sequence was...

Maintenance of Antibody to Pathogen Epitopes Generated by Segmental Gene Conversion Is Highly Dynamic during Long-Term Persistent Infection▿ †

Zhuang, Yan, Futse, James E., Brown, Wendy C., Brayton, Kelly A., Palmer, Guy H.

Multiple bacterial and protozoal pathogens utilize gene conversion to generate rapid intrahost antigenic variation. Both large- and small-genome pathogens expand the size of the variant pool via a...

Coinfection with Antigenically and Genetically Distinct Virulent Strains of Babesia bovis Is Maintained through All Phases of the Parasite Life Cycle▿

Berens, Shawn J., Brayton, Kelly A., McElwain, Terry F.

Antigenic polymorphism is a defining characteristic of the Babesia bovis variable merozoite surface antigen (VMSA) family. Sequence analysis strongly suggests that recombination between virulent...

Physical Linkage of Naturally Complexed Bacterial Outer Membrane Proteins Enhances Immunogenicity▿

Macmillan, Henriette, Norimine, Junzo, Brayton, Kelly A., Palmer, Guy H., Brown, Wendy C.

The outer membrane proteins (OMPs) of bacterial pathogens are essential for their growth and survival and especially for attachment and invasion of host cells. Since the outer membrane is the...

Composition of the Surface Proteome of Anaplasma marginale and Its Role in Protective Immunity Induced by Outer Membrane Immunization▿

Noh, Susan M., Brayton, Kelly A., Brown, Wendy C., Norimine, Junzo, Munske, Gerhard R., Davitt, Christine M., ...

Surface proteins of tick-borne, intracellular bacterial pathogens mediate functions essential for invasion and colonization. Consequently, the surface proteome of these organisms is specifically...

Superinfection as a driver of genomic diversification in antigenically variant pathogens

Futse, James E., Brayton, Kelly A., Dark, Michael J., Knowles, Donald P., Palmer, Guy H.

A new pathogen strain can penetrate an immune host population only if it can escape immunity generated against the original strain. This model is best understood with influenza viruses, in which...

Tick-Borne Transmission of Two Genetically Distinct Anaplasma marginale Strains following Superinfection of the Mammalian Reservoir Host▿

Leverich, Christina K., Palmer, Guy H., Knowles, Donald P., Brayton, Kelly A.

Strain superinfection affects the dynamics of epidemiological spread of pathogens through a host population. Superinfection has recently been shown to occur for two genetically distinct strains of...

Independence of Anaplasma marginale Strains with High and Low Transmission Efficiencies in the Tick Vector following Simultaneous Acquisition by Feeding on a Superinfected Mammalian Reservoir Host▿

Ueti, Massaro W., Knowles, Donald P., Brayton, Kelly A., Palmer, Guy H.

Strain superinfection occurs when a second pathogen strain infects a host already carrying a primary strain. Anaplasma marginale superinfection occurs when the second strain carries a variant...